The circuit has a -0.5v at vin, the source is fed back to the inverting input. fet is n channel d to 0v, inverting input connected to -5V via 10K to allow inverting input to be driven to same potential as vin i.e. -0,5v. the output is taken from the 1M, so -0.5/1M is the current o/p.
Note: vin...