Hi again, sorry for the slow reply.
Ideally I would like to see some good clear photos of the top and bottom side of the board with the large components removed. Then I think you should refit all the components, INCLUDING the 555 but NOT the micro and NOT the ULN2803s, and measure the voltages on all the marked points on this picture.
Attach your multimeter negative to the 0V point, which is on pin 1 of the 555 or anything connected to it. Measure voltage on each point, on the underside. Have the board powered up from 12VAC but with nothing else plugged into it.
I'm afraid all the letters are mirrored, because I annotated a picture that I had mirrored so that it would correspond to the top view picture. I'm sure you can handle it. The points are marked A~Q excluding I, L and O.
Measure the voltages on a good board and on the faulty board.
I'm sorry to put so much work on you. Normally a fault like this would be easy to find but there seems to be something strange about this board. CC and I are having a lot of trouble figuring out just how it's supposed to work and what's gone wrong.
Edit: You might want to clean all the crusty flux off the board before you take the photos. Use a cotton swab (a stick with cotton wool wound around each end - we call them 'cotton buds' over here) with isopropyl alcohol or any other solvent, and rub the board hard to get it really clean.